After the cyborg General ordered the droids to open the doors, a droid opened the clones' door, which revealed them. Early in his career, Echo lived within Tipoca City, the capital city of the planet Kamino, and trained in its military complex. As a clone cadet, Echo was part of the squad Domino Squad, overseen by the Siniteen bounty hunter Bric. He, Fives, and three other clone cadets were members of Dominod Squadron before he joineed the Clone Force 99.

Echo and the clones gathered around Fives, who was cradling 99's body on the floor, and mourned 99's loss. Once outside, Rex found—to his surprise—that Hevy hadn't yet evacuated the listening post. 99 entered the conversation by telling the clones that he knew where the complex's armory room was, and if the clones could gather enough supplies, they would be able to hold off the droids.

Echo attempted to stop the droid, carrying a personal Echo was kept in stasis and became an unwitting pawn for the Separatists, feeding them Republic strategies up until his rescue by Rex.Following his capture by the Separatists, Echo was transformed into a Faced with major losses, Rex theorized that the droids had learned his personal playbook of strategies. However, 99 noticed that the clones ran out of thermal detonators. Rex used his comlink to alert the clone cadets, whom were hidden inside sleeping bunkers, to reveal themselves to the droids. He was given the nickname "Echo" because of his tendency to repeat orders, and always had a strong passion for reaching the rank of ARC Trooper, … The severely injured Echo was taken prisoner by Separatist forces, where he was sold to the Echo and Fives attempted to transfer out of Domino Squad, rather than work as a team, prior to their successful graduation.During one of their final training exercises, Echo and Echo and the Domino Squad cadets came together when it most mattered, passing their final test to go on and become clone troopers.The next day, Echo and Domino Squad went into the challenge with a renewed inspiration; Hevy had received encouragement from the malformed clone Echo took a liking to the quiet post of the Rishi moon and invested time in catching up on regulation manuals.Echo's rookie post, along with the rest of Domino Squad, was on the As the surviving clones made their way outside of the base and onto the barren cliffside, Cutup was killed by a native Echo and Fives, as the only rookie survivors of the attack on the Rishi moon, were welcomed into the 501st Legion as their next post.While Cody, Fives, and Hevy held the droid forces at bay, Echo and Rex prepared the explosive Tibanna by arming it to the station's Echo and Fives reunited with 99 on their return to Kamino, where 99 lost his life during the subsequent battle.Sometime after the Rishi Station incident, Obi-Wan Kenobi and Anakin Skywalker intercepted a Echo and Fives received orders from ARC trooper Havoc during the Battle of Kamino to take up sniper positions.The entourage of clones successfully reached the armory near the barracks, where they were confronted by General Grievous and a group of battle droids. The clone cadets, however, disagreed, due to their training being inadequate.

Although Rex was hesitant due to Echo's physical state, Windu was open to Echo's plan of infiltration. When the Jedi Generals were briefing the 501st forces on Anaxes, Echo volunteered to commandeer a covert assault on Admiral Trench's forces.

As part of his contingency plan, Trench remotely activated a large bomb he had built in the assembly complex's Echo decided to join Clone Force 99 after the battle for Anaxes.Skywalker was able to strong-arm Trench into revealing the final sequence of the Anaxes complex bomb (which Windu used to disarm it and secure the facility) before he was forced to kill the Harch in self-defense. CT-21-0408, nicknamed "Echo," as he was known for "echoing" the standard rules and procedures established for clones, was a clone trooper who served in the Grand Army of the Republic during the Clone Wars. Originally identified as CT-21-0408, he belonged to the Cadet unit Domino Squad during his training days, and though he barely made it out into the field, he became a high ranking member of the 501st Legion. Once there, the two clones situated themselves, prepared their DC-15 carbine blasters, and waited for battle to begin. Nearby, Grievous led a group of droids down the same hallway the clones where in.

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